The USS Enterprise is sent to observe the "Class M" planet Nibiru, but finds a volcano on the verge of erupting and wiping out its primitive inhabitants. In the process of launching a dangerous mission to halt the eruption, first officer Spock's life is jeopardized, forcing Kirk to break the "Prime Directive" and reveal the Enterprise to the planet's civilisation to rescue him. As a consequence, a number of indigenous people begin to worship the ship as it leaves.
After being called back to Earth, Kirk is relieved of command, and Admiral Pike resumes command of the Enterprise. Shortly afterward, Starfleet agent John Harrison, with the compliance of a Starfleet officer, bombs a secret "Section 31" installation in London. An emergency meeting of high ranking officers is called at Starfleet headquarters, which Kirk attends as Pike's first officer. The meeting is attacked by a gunship piloted by Harrison. Kirk takes the gunship down, but Harrison escapes and Pike dies.
After Pike's funeral, Admiral Marcus (Weller) authorizes Kirk to hunt down Harrison, who has fled to the Klingon homeworld of Qo'noS ("Kronos"). Since Qo'noS lies deep in Klingon territory and Starfleet is on the brink of war with the Klingon Empire, the Enterprise is supplied with 72 long-range prototype photon torpedoes, and is ordered to fire them all at Harrison's location once he is found. Believing the torpedoes could be dangerous to the ship, chief engineer Montgomery Scott refuses to take them aboard and tenders his resignation when ordered to accept them. The Admiral's daughter, scientist Carol Marcus also joins the crew, under a false identity, and Pavel Chekov is promoted to Chief Engineer.
En route to the Klingon homeworld the Enterprise's warp core malfunctions, leaving the ship stranded in space. With repairs underway, Kirk, Spock and Uhura use a previously commandeered trader ship to enter Qo'noS. However, they are detected by Klingon patrol ships and are forced to land. Despite Uhura's attempts to negotiate, the Klingons prepare to kill the trio, when a mysterious figure attacks the Klingons. After wiping them out, the attacker reveals himself to be John Harrison. He confronts the landing party, but surrenders after learning the precise number of torpedoes aimed at him....
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